By Israel Zelitch
ISBN-10: 0127792503
ISBN-13: 9780127792507
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4. CAROTENOIDS The carotenoids, like the quinones of the chloroplast, are also made up of isoprene units, and the formula of the most common carotenoid, j3-carotene, shows that it is apparently made up of eight isoprenes by the head-to-head association of two units of four isoprenes (two diterpenes). ^-Carotene skeleton C H 2= C — C H = C H 2 Isoprene 36 /. Biochemical and Photochemical Aspects ι ' U ι ι— 400 500 ι ι : 600 Wavelength, nm ι 700 FIG. 2. The absorption spectra of chlorophylls a and b in methanol and of β-carotene in hexane (from French and Young, 1956).
The mRNA attaches to ribosomes and moves across ribosomes to bring successive nucleotide triplets (codons) into position to select the correct aminoacyl-tRNA precursors. A given mRNA molecule generally works simultaneously on many ribosomes (polyribosome), hence many codons function at a given moment. As the amino acids are polymerized into proteins on ribosomes, there is a concomitant release of tRNA. Initiation of protein synthesis occurs in ribosome subunits, which can be detected by their lower molecular weights on sedimentation in the ultracentrifuge.
Porter and Anderson, 1967). 2) / 2. Chloroplasts and Their Activities 37 Isopentenyl pyrophosphate is then converted to geranyl pyrophosphate (C 1 0), farnesyl pyrophosphate (C 1 5), and geranylgeranyl pyrophosphate (C 2 0). These C 2 0 compounds then condense head-to-head to produce carotenoid structures. A sequential desaturation pathway probably produces the various carotenes, and oxygenated carotenoids arise late in the biosynthetic pathway from carotenes. Although carotenoids do not usually absorb light directly in the photosynthetic process, carotenoids function to inhibit the photooxidation of chlorophyll.
Photosynthesis, Photorespiration, and Plant Productivity by Israel Zelitch
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